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in reply to Re^4: apache 2.4 mod_perl permission denied on standard modules
in thread apache 2.4 mod_perl permission denied on standard modules

This really sounds as if there is at least one entry in @INC which is not readable for the user Apache is running as. As soon as Perl encounters such a directory, it will stop searching. So you should make sure that the relevant directories you have in @INC appear before the bad ones where the user does not have read permissions.