If what's within the angle brackets is neither a filehandle nor a simple scalar variable containing a filehandle name, typeglob, or typeglob reference, it is interpreted as a filename pattern to be globbed, and either a list of filenames or the next filename in the list is returned, depending on context. This distinction is determined on syntactic grounds alone. That means "<$x>" is always a readline() from an indirect handle, but "<$hash{key}>" is always a glob(). That's because $x is a simple scalar variable, but "$hash{key}" is not--it's a hash element.