use strict;

my $HoA_ref = {'GAT' => [2, ['ttt',-3,1],['ttc',-3,3],['ccc',-1,2]],
               'AAA' => [13,['aaa',-1,2],['atg',-2,2]], 
               'TTT' => [11,['tta',-2,1],['atc',-3,3]]
               };


sub print_zHoA {
    my ($HoA) = shift;
    foreach my $mc (sort {$HoA->{$a}->[0] <=> $HoA->{$b}->[0]} keys %$HoA){
      print "$mc $HoA->{$mc}->[0]:\n";
      print join(',', map {$_->[0]} splice( @{$HoA->{$mc}}, 1)) . "\n";
    }
}

print_zHoA($HoA_ref);