use strict;
use warnings;

my @phrases=('(arf)71', 'p65(91)', 'H:223', 'Phospholipase A2 inhibitor', 'Phospholipase A2', 'Phospholipase', 'PLI-A', 'PLI-B', 'PLI');

my $string = join '|', map { quotemeta } @phrases;

my $in = "(arf)71 p65(91) H:223 Phospholipase A2 inhibitor ( PLI ) , purified from the blood plasma of the Habu snake ( Trimeresurus flavoviridis ) , was separated into two distinct subunits , PLI-A and PLI-B";

my $out = "#(arf)71# #p65(91)# #H:223# #Phospholipase A2 inhibitor# ( #PLI# ) , purified from the blood plasma of the Habu snake ( Trimeresurus flavoviridis ) , was separated into two distinct subunits , #PLI-A# and #PLI-B#";

#while($in =~ s/(^|\s)($string)(\s|$)/$1#$2#$3/) {};
while($in =~ s/(^|\s)($string)(\s|$)/$1#$2#$3/g) { pos() = pos() - 1; };

print "success!!\n" if($in eq $out);
print "in: $in\n";
print "out: $out\n";