#!/usr/bin/perl
use strict;
use warnings;

my $data = "Antler embedded in mound at South Street, Avebury, Wiltshire, England. Comment (lab): Collagen fraction used";

my @result;
while ($data =~ /\b([A-Z][a-z]*)/g) {
    push @result, $1;
}
print join(' ', @result), "\n";

##</code><code>##

The regular expression:

(?-imsx:\b([A-Z][a-z]*))

matches as follows:
  
NODE                     EXPLANATION
----------------------------------------------------------------------
(?-imsx:                 group, but do not capture (case-sensitive)
                         (with ^ and $ matching normally) (with . not
                         matching \n) (matching whitespace and #
                         normally):
----------------------------------------------------------------------
  \b                       the boundary between a word char (\w) and
                           something that is not a word char
----------------------------------------------------------------------
  (                        group and capture to \1:
----------------------------------------------------------------------
    [A-Z]                    any character of: 'A' to 'Z'
----------------------------------------------------------------------
    [a-z]*                   any character of: 'a' to 'z' (0 or more
                             times (matching the most amount
                             possible))
----------------------------------------------------------------------
  )                        end of \1
----------------------------------------------------------------------
)                        end of grouping
----------------------------------------------------------------------