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PA5098  PA5100  PA5092  PA3175

PA5098  PA5100  1
PA5098  PA5092  2
...
PA5100  PA5092  1
PA5100  PA3175  2
PA5092  PA3175  1

PA5098     PA5100    1
PA5098     PA5092    2
...
PA5100     PA5092    2
PA5100     PA3175    3
PA5092     PA3175 3

############# open pathways output list with PA01 locus IDs
my $in=$ARGV[1] || "pathways.col";
...
        }
        }
        }