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dddddddddBBBBBBBBBBBBDDDDDDDDBBBBBBBBBBBBBBddddddddddddddddddddBBBBBBB
+BBBBBBBDDBBBBBBBBddddddddddddd

use strict;
use warnings;
...
    print $seg."|".$seg_start."-".$seg_end."\n";
}