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  "A C D"

  /A(.*?)C(.*?)D/

  "ABICID"

  "A--C-D"

  if ($string =~ $test) {
    for my $offset (1 .. $#-) {
...
    # now extract only the matched extent
    push @hyph_padded_seq, substr($test, $-[0], $+[0] - $-[0]);
  }