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use Data::Dumper;

...
foreach my $aref (@aoa) {
    print "$aref->[0]\t\t $aref->[1]\t $aref->[2]\t $aref->[3]\n";
}

GAL_up800.fasta          6       800     4.8
hm01g.fasta              18      2008    36.144
hm02r.fasta              9       1000    9
train_muscle.fasta       7       50      0.35