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Before s///:     ,  ,  ,  ,  ,  ,  , 6
After 1st pass:  ,,|  ,  ,  ,  ,  , 6
After 2nd pass:  ,,  ,,|  ,  ,  , 6
After 3rd pass:  ,,  ,,  ,,|  , 6

1 while s{, +,}{,,};

s{, +(?=,)}{,}g;