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C:\test>774383.pl
Gene1:Gene2 tags::
...
         spcs  => ["spc6", "spc8", "spc6", "spc7"],
       },
}

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#! perl -s
use 5.010;
...
GeneE,GeneF,spc3,spc1
GeneX,GeneY,spc6,spc8
GeneX,GeneP,spc6,spc7