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for (grep $tree{$_}, keys %tree) {
   my @g = $_;
   push @g, $_ while $_=$tree{$_};
   print(join(':', @g), "\n");
}

for (grep $tree{$_}, keys %tree) {
   my $g = $_;
   $g .= ":$_" while $_=$tree{$_};
   print("$g\n");
}