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H P L A   143 TYR   202  OH  -->  O2   2014 MC9   500 A
H P L A   143 TYR   202  OH  <--  O2   2014 ASH   500 A
...
H P L A   237 SER   532  OG  <--  O1   2015 AGM   500 A
H P L A   274 ARG   821  NH1 -->  O1   2015 MC9   500 A
H P L A   278 SER   851  OG  -->  O2   2014 VIA   500 A

#!/usr/bin/perl -w

...
  while $fh->getline; 

$fh->close;