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Use of uninitialized value $1 in hash element at C:\Users\Jems\Desktop
+\Perl\test\test2script.plx line 26.
Use of uninitialized value $1 in hash element at C:\Users\Jems\Desktop
+\Perl\test\test2script.plx line 27.
Use of uninitialized value $1 in hash element at C:\Users\Jems\Desktop
+\Perl\test\test2script.plx line 28.

for ( read_file $uniprot ) {
   /(.{6})\s+.+=([^\s]+)/;
...
    push @activline, "$1 | $2 | $activ{$1}\n" if $activ{$1}; #26
    push @antioxline, "$1 | $2 | $antiox{$1}\n" if $antiox{$1}; #27
    push @toxinline, "$1 | $2 | $toxin{$1}\n" if $toxin{$1};#28

Q197F8 ORFNames=IIV3-002R 
Q197F7 ORFNames=IIV3-003L 
...
Q6GZX0 ORFNames=FV3-005R ;PF02393
Q91G88 ORFNames=IIV6-006L ;PF12299;PF04383
Q6GZW9 ORFNames=FV3-006R