Possibly it is related to the lstat call on line 442.

What happens if you add a stat call with the same arguments immediately after it? This will reset the value in _

...
$sub_nlink = (lstat ($no_chdir ? $dir_pref . $FN : $FN))[3];
my $debug = stat ($no_chdir ? $dir_pref . $FN : $FN))[3];

if (-d _) {
...
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