I've worked out the problem, in case anyone's interested - instead of...
$ftp->quot("user destination_username destination_password")
...do this...
$ftp->login("destination_username", "destination_password")
This will then allow you to ftp through a type 3 firewall (see perldoc Net::Config for definition of a type 3 firewall).
I suppose this was a bit obvious though...
Andrew
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