In your second example, the inconsistency between the button text and the sub is because of a difference in when the code is executed. When you call
$mw->Button, the code you have in your constructor call is run right then, so
"Button $_" is converted to a static string, and
Button saves a copy of that static string somewhere, so even if
$_ changes the button's text stays the same. But the anonymous
sub you give as the argument to
-command isn't executed until the button is clicked. By then the value of
$_ will probably have changed, and so the sub will use the current value of
$_ instead of its value when the constructor was called.
As others have said, your first example uses a closure, which implicitly creates a copy of the variable when the anonymous sub is created, so the sub refers back to that variable. Closures keep lexically-scoped variables (those created with my), but $_ isn't lexically scoped. That's the key to the difference between the two.
You could also solve the problem by forcing the anonymous sub to resolve $_ when it's created, using eval:
-command => eval "sub {print $_}"
(n.b. I haven't tested this and it might not be exactly right). But using closures is much easier, faster, and more elegant.
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