Update: He corrected himself when I was writting my post. Ignore this post.
No, the closing tag is optional (in HTML 4.01). Note the "O" for "Optional":
<!ELEMENT LINK - O EMPTY -- a media-independent link -->
<!ATTLIST LINK
%attrs; -- %coreattrs, %i18n, %events -
+-
charset %Charset; #IMPLIED -- char encoding of linked reso
+urce --
href %URI; #IMPLIED -- URI for linked resource --
hreflang %LanguageCode; #IMPLIED -- language code --
type %ContentType; #IMPLIED -- advisory content type --
rel %LinkTypes; #IMPLIED -- forward link types --
rev %LinkTypes; #IMPLIED -- reverse link types --
media %MediaDesc; #IMPLIED -- for rendering on these media
+ --
>
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