UNTESTED code but it should give you some ideas:
open A, '<', 'fileA' or die "Cannot open 'fileA' $!";
my %lines;
while ( <A> ) {
$lines{ $. } = () if /X/;
}
close A;
open B, '<', 'fileB' or die "Cannot open 'fileB' $!";
my $count;
while ( <B> ) {
s/(Y)/ exists $lines{ ++$count } ? 'Z' : $1 /eg;
print; # updated to print lines
}
close B;
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