It helps to know that the loop is expanded to:

while ($_ = <SOME_FILE_HANDLE>){
   ...
}

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This shows that <...> is nothing more than a loop condition: the loop body is executed immediately after each evaluation of the <...> operator. This is how a while loop works.

In reply to Re^3: Different behaviors between "while" and "map" by betterworld
in thread Different behaviors between "while" and "map" by lightoverhead

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