Sounds like mencoder's behaviouris dependent on whether it's connected to a terminal or not. If so, you want to look a pseudo ttys. I'd look at IO::Pty, possibly called via IO::Run.
By the way,
$arg1 =~ s/ /\\ /g;
$arg2 =~ s/ /\\ /g;
is very badly done. Use the multiple argument form to avoid having to quote at all.
open(CONVERSION, '-|',
mencoder => $arg1,
-oac => 'mp3lame',
-aid => 128,
-ovc => 'xvid',
-xvidencopts => 'bitrate=-1',
-vf => 'scale',
-zoom,
-xy => 320,
-o => $arg2,
);
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