From open:

If the filename begins with '|' , the filename is interpreted as a command to which output is to be piped, and if the filename ends with a '|' , the filename is interpreted as a command which pipes output to us. See "Using open() for IPC" in perlipc for more examples of this. (You are not allowed to open to a command that pipes both in and out, but see IPC::Open2, IPC::Open3, and "Bidirectional Communication with Another Process" in perlipc for alternatives.)

(emphasis mine)