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Starting your script like: perl -d scriptname.pl will bring it up in the perl debugger. Read perldebug. You may also want to dump your data structures at opportune moments with Data::Dumper.
Your code is well formatted and except for the while() loop surrounding a bunch of subroutine definitions ( the subroutine definitions should be outside the execution path ) otherwise the code is very readable. The parameter passing is incorrect however. Subroutine calls like some_sub( %hash ) are usually not correct as perl passes your hash structure as a flat list which then becomes a local copy in the subroutine. What you probably should be doing is passing a reference to the hash. See: perlreftut. This is why your data structure is not getting updated as your routines only modify their local copy of the structure.
s//----->\t/;$~="JAPH";s//\r<$~~/;{s|~$~-|-~$~|||s
|-$~~|$~~-|||s,<$~~,<~$~,,s,~$~>,$~~>,,
$|=1,select$,,$,,$,,1e-1;print;redo}
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