Could also say: 
my $rc = reverse uc($dna) =~ tr/ATGC/TUCG/r;

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Laurent_R said Except that the OP said: without using s// or tr///. That's part of the assignment.

Yes, my bad.

In reply to Re^2: Trying to use substring to find a reverse complement in DNA, by Cristoforo
in thread Trying to use substring to find a reverse complement in DNA, by tryingnottofailmytes

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