m/IDX/ returns true (1) if $_ matches the pattern and false otherwise. ~ is the bitwise not operator (aka one's complement - see
Symbolic Unary Operators). It inverts all the bits in a number.
Your match doesn't match $_ and so returns false, which is complemented to 0xffffffff, which, in decimal, prints as 4294967295.
Try
perl -e 'print ~0'
or, for 64-bit perl:
perl -e 'print ~0 & 0xffffffff'
Update: Added code examples & link to ~ in perlop
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