The upload file filesystem browser widget is a function of the OS, not the browser, isn't that odd? It gives the browser a way to upload the file, but afaik, it does not mention the path in the POST. So, I think the answer is going to be no: there's no way to let the user browse their file system from your CGI and POST a directory -- without some kind of magic (ie, non CGI.pm) hackery anyway.
UPDATE: "widget is a function of the OS, not the browser" / "No, it's not. It is completely application dependent." Well, the applications I mean are firefox and IE, which load the OS widget, which is why css doesn't work on the file upload buttons. "path in the POST" / "some user agents" So? It's not really something you can depend on then, is it? I think that's not only application dependent, but OS dependent.
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As stated by others (and the documentation), the path is not dependably returned by the user's browser. So, if you're really lucky, you might be able to find this in $filename = param('uploaded_file');. Of course, this assumes that the user actually uploads a file. Probably not what you want but it is a faint possibility.
What I've done to get around a similar problem is to have (intranet) users start the process with a local program (VBS in my case) which gets the required info from the local system and opens an internal webpage with the appropriate parameters to do the work.
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